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Call of Duty: Ghosts is set in a near future that follows the nuclear destruction of the Middle East. The oil-producing nations of South America form the "Federation of the Americas" in response to the ensuing global economic crisis and quickly grow into a global superpower, swiftly invading and conquering Central America, the Caribbean, and Mexico.
The game's main protagonists are the Ghosts, a force of U.S. Special Operations personnel trained to conduct clandestine missions behind enemy lines. The unit is led by retired U.S. Army Captain Elias Walker (Stephen Lang). He is joined by his sons Logan and David "Hesh" Walker (Brandon Routh), along with a trained German Shepherd named Riley, Captain Thomas A. Merrick (Jeffrey Pierce), and Sergeant Keegan P. Russ (Brian Bloom).
The game's main antagonist is Gabriel Rorke (Kevin Gage), the former leader of the Ghosts who works for the Federation after being captured and brainwashed by the Federation via torture and hallucinogens.
In 2017, U.S. Army officer Elias Walker tells his teenage sons Hesh and Logan the legend of how the Ghosts, an elite coalition of all U.S. Special Operations units, first appeared. Meanwhile, in space, the "Federation of the Americas" hijack the Orbital Defense Initiative (ODIN), an orbital superweapon that consists of a network of satellites that utilize kinetic bombardment. The Federation use it to destroy several cities in the southwestern United States. U.S. astronauts Baker and Mosley sacrifice themselves to self-destruct the space station and prevent ODIN from launching other payloads further inland. The Walker family narrowly escapes the destruction of San Diego, California.
Ten years later in 2027, the war between the United States and the Federation has been a bloody stalemate along a front that comprises the destroyed cities (known as "No Man's Land"), which has become a war of attrition as more Federation forces attempt to break the frontlines. However, the U.S. stands it ground, fighting near the former United States-Mexico border. Hesh and Logan are part of a U.S. special unit under the command of Elias and are accompanied by Riley, their loyal military-trained German Shepherd. During a mission into No Man's Land, they spot American Federation agent Gabriel Rorke interrogating Ghost member Ajax. The brothers meet and join Ghost members Thomas Merrick and Keegan Russ in their mission to rescue Ajax, but are unsuccessful as he is killed.
The brothers return to Santa Monica, California, where U.S. forces hold out against an assault from the Federation, and reunite with their father, who reveals himself as the leader of the Ghosts. Recruiting his sons into the Ghosts, Elias tells them about how Rorke was the Ghosts' previous commanding officer until Elias was forced to abandon him during a successful mission in Caracas to assassinate former Federation leader, Diego Almagro. The Federation captured and brainwashed Rorke, and are now using him to hunt down the Ghosts.
The Ghosts assault Rorke's base and capture him, but Federation forces destroy their plane and rescue him. The team lands in the Yucatán Peninsula, where they witness the launch of a mysterious Federation missile. The Ghosts infiltrate a Federation base and discover plans to create a new superweapon. Constructing a counter-attack, the Ghosts make significant gains against the Federation, successfully destroying their Atlas oil platform in Antarctica and sinking a destroyer guarding the Brazilian coast. The team then breaks into a Federation laboratory and discovers that the Federation reverse-engineered ODIN into their own orbital bombardment system named LOKI.
After destroying a Federation factory in Rio de Janeiro, the Ghosts regroup at a safehouse in Las Vegas, but are captured by Rorke, who executes Elias. In a last-ditch effort, the U.S. pools all its remaining forces in a synchronized all-out assault. A team of soldiers board a shuttle and take over the Federation space center in the Atacama Desert, while forces in space take over the Federation space station. They seize control of LOKI and decimate Federation forces by turning its weapons on them. Hesh and Logan then pursue Rorke to avenge their father, disobeying Merrick's orders. Despite being shot by Logan and left to drown, Rorke survives the encounter and captures Logan, aiming to try and brainwash him into becoming a fellow Federation agent.
In a post-credits scene, Logan is seen being kept inside a pit in the jungle, presumably going through the same torture methods that Rorke went through.
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